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muricatus, and show that ovotestes occur at frequencies consistent with a mode of **** determination that is intermediate between GSD and TSD. This is the first evidence suggestive of underlying unidentified **** determining loci in a species that has long been used as a model for TSD.Osteostraci and Galeaspida are stem-gnathostomes, occupying a key phylogenetic position for resolving the nature of the jawless ancestor from which jawed vertebrates evolved more than 400 million years ago. Both groups are characterized by the presence of rigid
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