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Coevolution of the gene encoding the T4SS system is a possibility indicated by our research concerning Bradyrhizobium and Mesorhizobium. In the Bradyrhizobium species, the copG, traG, and virD2 gene cluster is uniquely configured. Gene deletion and complementation experiments on T4SS1 demonstrated its positive role in nodulation. Accordingly, the T4SS system likely plays a critical role as a determinant for sym
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