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Vegetative parts of bilberry (Vaccinium myrtillus) are important forage for many boreal forest mammal, bird and insect species. Plant palatability to insects is affected by concentration of nutrients and defense compounds in plants. We expected that palatability of bilberry leaves to insect herbivores is influenced by light availability and soil productivity (both affecting nitrogen concentration and constitutive carbon-based defense compound concentration) and herbivory by mammals (affecting nitrogen concentration and induced carbon-ba
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